Altogether 50 different alleles were recorded in populations situated South of the Holdhaus line, whereas only 23 different alleles were found in populations North of it. This is also reflected by the allele numbers corrected for the number of investigated populations (rarefied to 34 populations, Krebs 1999): The populations situated South of the Holdhaus line show 41.5 alleles, whereas the populations North of it have only 22.0 different alleles.
With the exception of the seven private alleles in the Black Forest, the Vosges, and parts of the Swiss and French Jura (see above), all alleles that were found in populations North of the Holdhaus line are also found in populations located South of it. The Northern populations are, therefore, a section of the genetic richness of the Southern populations.
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