expected frequency of AB gametes after one generation of natural selection and recombination is

W11X2 + W12X1X2 + W13X1X3

which is directly comparable with equation 6.21 for one allele at a diallelic locus. This can be simplified by expanding the (1 - r)w14x1x4term:

wnx 1 + ww 12X1X2 + ww 13X1X3 + ww 14X1X4 rw^xx + rw2 3X2X3

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