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which after addition simplifies to

as the expected genotypic mean value of all individuals with an M1/M1 marker locus genotype.

Since j/4 of all individuals in the F2 population are expected to have an M1/M1 marker genotype by Hardy-Weinberg, we can multiply by a factor of 4 to obtain the actual genotypic mean of that subset of individuals in the F2 population with M1/M1 genotypes:

The same logic can be used to obtain expressions for Gpop and G for the M1/M2 and M2/M2 marker genotypes. Note that environmental variation in phenotype does not need to be accounted for because

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