where PI(t) indicates the probability that the nucleo-tides at a given site are identical. The exponential term is now e-8at because there are two DNA sequences that can change independently so that the chance of the nucleotide remaining the same decreases twice as fast with time. The probability that two sites are different or divergent - call it d - over time is one minus the probability that the sites are identical, so that which is analogous to an exponential growth equation for a population with a carrying capacity. As d - 3(1 - e-8at) 4

A3 1

-Initially G

A3 1

0 0

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