genotype a value of -a. The expected frequency of each of the three genotypic values within the M1/M1 marker-locus genotype is then the product of the genotype frequency and the corresponding genotypic value.

The portion of the mean genotypic value for the entire F2 population due to those individuals with an M1/M1 marker genotype, call it GMpm , is the sum of the genotype-frequency-weighted genotypic values for each 0TL genotype associated with an M1/M1 marker genotype:

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