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e is also the probability that the alleles in a genotype are identical by descent or are autozygous. The probability that two alleles in a genotype are not identical by descent or are allozygous is then 1--. Through

2Ne the process of random sampling, populations can accumulate autozygous genotypes, as do populations where mating takes place among relatives.

We can use the probability of autozygosity in a finite population to define the fixation index (recall that it was defined as expected heterozygosity minus observed heterozygosity all over expected hetero-zygosity in Chapter 2) as

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