Varp pq

where the variance in allele frequency among n

1 n subpopulations is var(p) = — } (p. - p)2 and there n ~ 1 i=i are a very large number of subpopulations (Wright 1943a). If there is more variance in allele frequency then subpopulations differ more in allele frequency and the resulting FST is larger. For example, in Fig. 4.7 the average allele frequency or p in both sets of six subpopulations is 0.5. On the left the vari. . 3((0 - 0.5)2) + 3((1 - 0.5)2)

ance in pis-= 0.25 while

0 0

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