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This node defines the last common ancestor between the anamensis group and other hominins, defined by three derived features: the mandibular mental foramen opens laterally; molar enamel has increased in thickness (which is related to the next feature in that cusps and cristae are inflated); and the deciduous molar mesial marginal ridge is slight, with protoconid set anterior. The anamensis hominid is defined by only one apomorphy, the mandibular inferior torus is strongly developed; the lack of additional apo-morphies is clearly related to low specimen numbers and poor preservation.

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