The last common ancestor to H. ergaster and H. sapiens has a variable glenoid fossa depth. The vaginal process of the tympanic is large. There is a continued increase in cranial capacity, and the frequency of an occipito-marginal sinus has also increased. The orbital fissure is comma shaped. There is a continued reduction in palate prognathism, and the nasal clivus and nasal floor are defined by a smooth transition with overlap and a corresponding increased development of the incisive canal. There is also continued reduction in male canine size and in premolar size, even greater than the reduction in the molars.

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