Acknowledgements

This project was supported by the National Science Foundation (Grant SBR 9224151), Chicago Zoological Society, Sigma XI, Conservation International, and Berry College. We wish to thank the Ministry of Higher Education and the Department of Water and Forests for the opportunity to conduct research at Mantadia National Park, Madagascar. Frances White and Ken Glander provided important guidance while Rob Faigen, Ulrike Devoto, and Becca Lewis were invaluable as field assistants. This project could not have been completed without the assistance of head guide Ndrina Kotonirina. We thank Ellen Dierenfeld for nutritional analysis, and Amanda Wallace and Hoyt Hughie for laboratory assistance. We also thank two anonymous reviewers for helpful comments on the manuscript.

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