Thanks are due to S. Arrigo-Nelson, M. Banks, C. Borries, L. Godfrey, S. King, R. Lewis, T.L. Morelli, S. Nash, D. Overdorff, E. Patel, J. Powzyk, K. Samonds, and P. Wright for discussions and access to unpublished results which greatly improved this manuscript. The author's research at Tsinjoarivo was funded by the Margot Marsh Biodiversity Foundation, Primate Conservation, Inc., International Primate Protection League, and Stony Brook University; research facilitation was provided by ICTE (Stony Brook) and MICET (Madagascar).

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