For insightful comments that improved earlier versions of this manuscript, I thank Bob Sussman, Michelle Sauther, and two anonymous external reviewers. The following individuals and institutions greatly facilitated this project: the late Madame Berthe Rakotosamimana (University of Antananarivo), the Missouri Botanical Garden, the Peregrine Fund, the Masoala Integrated Conservation and Development Project, and my field assistants, Velo Norbert, Behasy Remy, and Moi'sse. The Tripartite Committee of the Malagasy Government gave permission to complete this research. Funding was provided by the Wenner-Gren Foundation, the Leakey Foundation, the National Science Foundation, Primate Conservation Inc., the Boise Fund, and Sigma Xi.

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