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a AGR, allo-grooming; MGR, mutual-grooming. Rates did not vary by category significantly.

a AGR, allo-grooming; MGR, mutual-grooming. Rates did not vary by category significantly.

allo-groom) also were similar to each other within food scarcity and food availability periods but both rates dropped significantly during food scarcity (MGR: X2=5.07, p<0.04, df =1; AGR: X2=3.67, p<0.05, df =1, Figure 2).

The directionality of grooming patterns did not vary seasonally. The male, female, or offspring were equally likely to initiate mutual-grooming bouts with each other and the adult female was least likely to initiate allo-grooming with the adult male (%2 = 10.23, p<0.03, df =4, Figure 3). Compared with social grooming, self-grooming rates were almost three times higher during food availability but also dropped to similarly low levels during food scarcity (%2 = 28.45, p <0.0001, df = 1, Figure 2).

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