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Figure 1. Observation grid. It combined existing permanent trails and small trails established for this study with location of randomly chosen forest plots of 5 m x 5 m used for characterisation of forest. Larger permanent trails (south-north) are indicated with a Roman numeral, east-west with a letter. Small trails divided the forest into quadrats of 10 m x 10 m, each quadrat unambiguously addressed by an Arabic number (south-north) and a letter (west-east), as on a chessboard. For subsequent analyses, locations were transformed into an xy-coordinate system.

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