a | = Malagasy lemurs. bData from Godfrey et al. (2005).

eSee Godfrey et al. (2005) for a review of calculating R.E.T.

(at least some time in the past), as seen in its extreme condition in extinct archae-olemurids (e.g., Godfrey et al., 2005, this volume).

Mammalian teeth, in addition to providing a record of both growth and development and evolutionary relationships (e.g., Schwartz and Dean, 2000), also reflect an individual's life experience or life story (e.g., Morbeck, 1997). Even among humans, diet and behavior leave a record of life on the teeth (e.g., Molnar, 1971). Despite the hardness of dental enamel (e.g., Maas and Dumont, 1999) and its assumed resistance to damage and crack propagation, tooth wear is common across the mammalian radiation (e.g., Hillson, 1986, 2005). Tooth damage, including severe wear, breakage, and pathology, is also common throughout the primate order (e.g., Schultz, 1935). However, to date data on patterns of dental health in lemurs are limited when compared to anthropoid primates, especially hominoids (e.g., Kilgore, 1989; Lovell, 1990). Patterns of dental damage often correlate with behavior in anthropoids, for example the high frequency of tooth loss resulting from breakage among male howler monkeys, which is often a result of intermale aggression (Smith et al., 1977). A similar pattern likely exists in

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