remained stable in location over several years as shown by the small shift of home range centers (Figure 10). Differences in home range size are most likely related to observation intensity, the range varying in the Lepilemur male L1m between 0.71 and 1.10 ha and 0.57 and 0.94 ha in the female L1f (Table 3). In addition, Lepilemur show differences in travel distances according to season (see below). Hence, range size also depends on the season when data are collected. A size around 1 ha may be a reasonable estimate for Lepilemur edwardsi in Ampijoroa, both male and female.

Nightly Path Length (Figure 11, Appendix A-VIII). There is no significant difference either across species or individuals (Kruskal-Wallis test, ns) if the entire year is considered. However, there is a conspicuous difference in the distribution of NPL over the year between Avahi and Lepilemur as shown in Figure 11. Towards the beginning of the rainy season in November/December, there is a major peak in path lengths in sportive lemurs, both male and female. During the rest of the year sportive lemurs have on average a lower NPL than do woolly lemurs.

Figure 8. Home ranges and range centers (harmonic mean) of the focal Avahi male based on all direct observations and sleep tree localizations for several consecutive years.
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