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a 100 x (optic foramen area/orbital area); SD, standard deviation.

a 100 x (optic foramen area/orbital area); SD, standard deviation.

lemurs (family Megaladapidae), Megaladapis (Peloriadapis) edwardsi (~ 90 kg) displays values lower than Megaladapis (Megaladapis) grandidieri (~75 kg) and M. madagascariensis(~ 45 kg). The largest-bodied species actually resemble living nocturnal lemurs in estimated degree of summation more so than their diurnal or cathemeral counterparts.

Overall, however, and despite their much larger body sizes, most extinct lemurs have indices well within the observed ranges of extant diurnal lemurs. In size-matched comparisons, all subfossil lemurs have OFIs lower than diurnal anthropoids (e.g., compare Archaeoindris to gorilla or Megaladapis to orangutans), and it seems reasonable to conclude that their visual acuity was also correspondingly lower, even if they were predominantly diurnal in their activity cycles. In other words, despite their large, "anthropoidlike" body sizes, the giant lemurs appear to be typical strepsirrhines in terms of their lower visual acuity. This conclusion needs to be qualified somewhat by the well-known observation that absolutely larger eyes, such as those of giant lemurs, may improve acuity to some unknown degree. This follows because photoreceptor cell size is more or less constant across animals of all body sizes (Land and Nilsson, 2002); a larger eye can therefore pack in an absolutely greater number of photoreceptors.

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