Because of space limitations, we cite review papers rather than reports of the original works. We apologise to the authors and warmly advocate the readers to go and read the original papers referenced in the reviews. We gratefully acknowledge Dominique Anxolabéhère, Romain Derelle, Alexandre Hassanin, Michael Manuel, Danielle Nouaud, Hadi Quesneville and Stéphane Ronsseray for discussions. We thank Peter Holland very much for insightful comments on a previous version of this manuscript, which led us to undertake new sequence analyses. We are grateful to Alessandro Minelli and Giuseppe Fusco for inviting one of us ( J.D.) to the evo-devo meeting held in Venice in May 2006 and for giving us the opportunity to publish our hypothesis in the present volume.

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