The authors thank the animal care staff of San Diego Zoo, the Smithsonian's National Zoological Park, Zoo Atlanta, Adventure World,

Zoologico de Chapultepec and the China Conservation and Research Centre for the Giant Panda for assisting in sample collection and recordkeeping. We are especially grateful to Desheng Li for technical assistance. We thank Lesley Northrop, Edy MacDonald, Angeles Pintada, Alan Fetter, Jean-Pierre Montagne, Kendall Mashburn and Nicole Presley. Funding was provided by the Alice C. Tyler Perpetual Trust, Friends of the National Zoo, the Zoological Society of San Diego and the Smithsonian's National Zoological Park.

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