Ammonoid Assemblages

Clymeniids dominate all Middle-Late Famennian ammonoid assemblages of the Uralian-Kazakhstanian Region. The clymeniid faunas include two suborders, four superfamilies, 14 families, 41 genera, and some 130 species. Most taxa (over 70%) belong to the suborder Clymeniina, with Cyrtoclymeniidae, Rectoclymeniidae, and Cymaclymeniidae dominant in diversity and number of individuals (Fig. 15.5) (Bogoslovsky, 1981; Nikolaeva and Bogoslovsky, 2005a).

Four successive assemblages corresponding to ammonoid genozones are recognized in the Uralian-Kazakhstanian Region: Cheiloceras, Prolobites-Platyclymenia, Clymenia-Gonioclymenia, and Kalloclymenia-Wocklumeria (Bogoslovsky, 1971, 1981) (Fig. 15.2). The succession of species within these genozones is similar to that recognized in Germany and Morocco and internationally (Korn, 1999; Becker and House, 2000; Becker et al., 2002), although there are regional differences. Because the ammonoid succession in Germany has been studied in great detail, the species-based zones there are referred to for comparisons with distant regions of the world (see Korn, 1999; Becker, 2000; Becker et al., 2002; Korn and Klug, 2002). The comparisons are closest at the supraspecific level but there are clear regional and local differences at the species level.

Famennian

Stage

Prolobites-Platyclymenia

Clymenia-Gonioclymenia

V

Kalloclymenia-Wocklumeria 0

-A

-B

n

0 A

0 B

| VI-C-11

| VI-C-2 |

VI-D-1

0 2

0

V0 F

Rhenish Massif and Morocco ss ge totyclymnia pompeckii— Platychmenia tenuis — Platyclymenia pl atyclymeniaplacida ••• — Platyclymenia fransita c Platyclymenia tschernyschewi — Platyclymenia subnautilina — Platyclymenia annulata — Platyclymenia gemma — Platyclymenia inosfranzevi= Trigonoclymenia spinosa -Trigonoclymenia sherubensis _ Trigonoclymenia tigra Trigonoclymenia orientals— Pleuroclymenia costata— Pleuroclymenia kasachstanica— Pleuroclymenia sp

Cyrtoclymenia involute/—

Cyrtoclymenia uralica — Cyrtoclymenia frechi —

_ Spinoclymenia aculeata—

Cyrtoclymenia planistriata Protactoclymenia m

C>rtoclymenia angustiseptata •

Protactoclymenia cara Protactoclymenia krasnopolski Pricella lanx Pricella stuckenbergi Pricella glabra

Sg1 o f sa

Pricella kajraktensis = Rectoclymenia roemeri—

Rcoome ahaaua ~

Rectoclymenia lenticella -Rectoclymenia luclulenta —

Rectoclymenia gracilis

Rectoclymenia fraudulen ta —

Rectoclymenia subplicata

Rectoclymenia jrnitima,

Rectoclymenia lyrata Rectoclymenia semilyrata Rectoclymenia tecta Falciclymenia.i

Faladymenia exceUens —

Falciclymeniamirabilis Falciclymenia plicata Falciclymenia uralica

Cteroclymenia rozmanae — Cteroclymenia infrequens _ Carinoclymenia beuelense Acriclymenia thaumasta—

Karaclymenia sperata— Aktuboclymenia ancestralis

Pinacoclymenia inexpectata

Clymenia aspis Clymenia laevigata

Clymenia cranoideformis c

Protoxyclymenia carinata

Protoxyclymenia dubia Protoxyclymenia intermedia -

Protoxyclymenia pseudoserpentina

Protoxyclymenia plana Kosmoclymenia (K) lamellistriata—

Genuclymenia aktubensis Genu clymen ia angelini ■ Genuclymenia frechi Genuclymenia karpinskii

Genuclymenia aktubensis • Genuclymenia sp.

Kosmoclymenia (K)inaequistriata — ■ Kosmoclymenia (K)timida

Kosmoclymenia - Kosmoclymenia (M)levis " Laminoclymenia aktubensis Laminoclymenia kiensis s o

C pa

Kazakhoclymenia medoevi -Cymaclymenia costata

Lissoclymenia wocklumeri ■

Laganoclymenialagana

Ornatoclymenia ornata -

Cymaclymeniastriata Cymaclymenia decorata Cymaclymenia fabra Cymaclymenia crenata Cymaclymenia barbarae Cymaclymenia subcompressa Cymaclymenia pseudocompressa

Cymaclymenia tuberculata Cymaclymenia

Cymaclymenia involvens

Genozones

Fig. 15.5 Distribution of the suborder Clymeniina in the Famennian of the Uralian-Kazakhstanian Region. This clymeniid suborder was the dominant component of the ammonoid faunas in this region. Solid black lines show the taxa recorded by Nikolaeva and Bogoslovsky (2005a). White lines are drawn based on records by other authors.

In the South Urals and Kazakhstan, recognition of species zones has always been very problematic because the sections are highly condensed (Bogoslovsky, 1971; Simakov et al., 1983). However, it is evident that the equivalents of some species zones recognized in Germany can be recognized there too, although the precise location of the zonal boundaries will require more data. The zonal boundaries in some sections are drawn tentatively and are the subject of ongoing research.

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