Patterns of Embryonic Development in Early to Middle Devonian Ammonoids

Susan M. Klofak,1 Neil H. Landman,2 and Royal H. Mapes3

'Division of Paleontology (Invertebrates), American Museum of Natural History, 79th Street and Central Park West, New York, NY 10024, USA, and Department of Biology, City College of the City University of New York, Convent Avenue and 138th Street, New York, NY, 10031, USA, [email protected];

2Division of Paleontology (Invertebrates), American Museum of Natural History, 79th Street and Central Park West, New York, NY 10024, USA, [email protected];

3Department of Geological Sciences, 316 Clippinger Laboratories, Ohio University, Athens, OH 45701-2979, USA, [email protected]

1 Introduction 15

2 Material and Methods 19

3 Results 20

3.1 External Features of the Ammonitella 20

3.2 Lirae Spacing 22

3.2.1 Mimagoniatitidae 23

3.2.2 Anarcestidae 25

3.2.3 Agoniatitidae 30

4 Discussion 30

5 Conclusions 35

Acknowledgments 36

Appendix 36

References 53

Keywords: Devonian, ammonitella, Agoniatitidae, Mimagoniatitidae, Anarcestidae, New York, Morocco

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