The author is grateful to Drs Chi-Hua Chiu and C.O. Lovejoy for insightful discussions on many of the topics covered here and thanks M. Ravosa and M. Dagosto for the opportunity to present this and other contributions directed towards understanding primate evolution. Ms Linda Gordon and Drs J. Mead and A. Gardner provided curatorial assistance at the Smithsonian Institution, Mr B. Stanley at the Field Museum of Natural History, and Mr L. Jellema at the Cleveland Museum of Natural History. Dr G. Gunnell, University of Michigan, generously shared unpublished data on hand and foot proportions in Plesiadapis cookei and J. Bloch and D. Boyer provided the opportunity to examine undescribed specimens of Phenacolemur.

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